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Scientific Name:
Humulus lupulus
Family Name:
Common Name:
hop, hops
Other Information
Cultivation, Conservation & Ecology
An article reviews recent studies on the response of hop (Humulus lupulus) to abiotic stresses in the context of the expansion of hop cultivation to semi-arid areas of Spain and Italy, providing an insight into the potential effects of the climate on the crop quality and production. Marceddu 2022
An article provides a review of available studies on the hop cyst nematode (Heterodera humuli). Darling 2022
A study presents a collection of 1592 images of the leaves from 12 hop (Humulus lupulus) varieties popular in southeastern Brazil with the purpose of developing pattern recognition and machine learning techniques for the classification of hop varieties. Castro 2021
A study evaluated whether use of copper-based fungicides negatively affects the antioxidant capacity of hops (Humulus lupulus) ethanol extracts. Chrisfield 2021
A study analyzed the contents of phthalic acid esters in the agricultural soil taken from a hop field in the Czech Republic. Langová 2020
An experimental study questions long-held notions regarding the effects of chilling and dormancy on flowering in hops (Humulus lupulus), suggesting hop production can be distributed more widely and may achieve four crops per year. Bauerle 2019
An article reports the dataset related to the genetic determination of Verticillium nonalfalfae infection of hop (Humulus lupulus). Svara 2019
A single application of a nonselective insecticide disrupted the biological control of the twospotted spider mite (Tetranychus urticae) in hops, resulting in a severe outbreak. Within the year, the population of resident predatory arthropods returned to normal. Iskra 2019
A survey of hop producers in Oregon and Washington states regarding cooperation in powdery mildew disease management suggested small-scale interactions being the most effective. Sherman 2019
Nitrogen fertilization was linearly related to the abundance of hop looper, Hypena humuli, which can cause substantial crop damage in hops, with a 2.5 increase in hop looper-days per kilogram of nitrogen fertilizer applied. Iskra 2018
A robust reverse transcription-recombinase polymerase amplification assay was developed for the analysis of hop samples for the presence of Hop stunt viroid. Kappagantu 2017
Incorporation of ground covers (red clover, or common yarrow/beebalm/red clover/annual sunflower) provided benefits for arthropod pest control in hop yards in Vermont, USA. Calderwood 2017
Bacterial endophytes from the stems of Humulus lupulus and 12 other plants grown in Poland were collected and characterized. Goryluk-Salmonowicz 2016
The microRNA response in Humulus lupulus to the recently discovered Citrus bark cracking viroid, considered one of the most devastating infections in hop, was studied. Mishra 2016
Novel virulence-associated proteins secreted to xylem by Verticillium nonalfalfae during colonization of hop plants were identified. Flajsman 2016
Complete mitochondrial genome of Verticillium nonalfalfae, the fungal pathogen causing wilt disease in hops, was sequenced. Jelen 2016
Delayed harvest was shown to increase the levels of linalool, geraniol, and myrcene, which contribute to the beer aroma, in hops of the Saaz variety, grown in the Saaz region of Czech Republic. Matsui 2016
The physiological and molecular biological response of hop plants to draught stress was studied in hop cultivars Aurora and Savinjski Golding. Kolenc 2016
An analytical model, estimating the influence of soil concentrations and other soil factors on the uptake of metals (Cd, Cr, Cu, Mo, Ni, Pb, and Zn) by hop crops, was developed. Novotná 2015
A meta-analysis of studies on the relationship between concentrations of microbial agents in the air of hop-handling farms and the occurrence of work-related respiratory and general symptoms in exposed workers was conducted. Mackiewicz 2015
Genetic mechanisms of resistance to a highly virulent strain of Verticillium albo-atrum, currently presenting a serious threat to hop production in Europe, were studied in resistant and susceptible hop plants. Cregeen 2015
Infection of hop cones by powdery mildew is most severe if inoculation occurs within 15-21 days after bloom, and fungicide effectiveness is similar when the applications are made season-long or targeted only to the juvenile stages of cone development. Twomey 2015
Differential expression of proteins responsible for the lethal vs. mild effects of Verticillium albo-atrum, a vascular wilt pathogen, in hops was investigated. Mandelc 2015
The occurrence, type and distribution of calcium oxalate crystals in the leaves and stems of Humulus lupulus, as well as several other plants, were characterized. Crystal sands were observed only in the pith tissue of Humulus lupulus. Tütüncü Konyar 2014
Development of biological control of a spider mite, Tetranychus urticae, and hop aphid, Phorodon humuli, in Oregon hop yards over a 9 year period was studied. While the potential of the biological control of the hop aphid is limited, it may be successful in case of the spider mites. Woods 2014
Pyraclostrobin, a fungicide used against powdery mildew, suppresses hop looper, Hypena humuli, a reemergent pest of hop. Woods 2014
Additive genetic parameters for yield, plant growth and cone chemical traits in hop were investigated. McAdam 2014
The molecular diversity of four hop stunt viroid variants was studied. Elleuch 2013
No significant infection-specific changes were identified in the proteomic analysis of incompatible response of hop root tissues to Verticillium albo-atrum, the causal agent of Verticillium wilt, a serious hop disease. Mandelc 2013
Reference genes for the development of Verticillium wilt-resistant hop cultivars were identified. Štajner 2013
Tetraploidization of the hop cultivar 'Sybilla' increased cone weight and the proportion of humulene, caryophyllene and farnesene, but reduced total essential oil yield of the plants. Trojak-Goluch 2013
Genetic transformation of hop plants with PAP1/AtMYB75 factor from Arabidopsis thaliana changed the color of the flowers and increased the contents of anthocyanins, rutin, isoquercitin, kaempferol-glucoside, kaempferol-glucoside-malonate, desmethylxanthohumol, xanthohumol, and α- and β-acids. Gatica-Arias 2012
Complete genomic sequence of American hop latent virus (AHLV) was determined. Eastwell 2012
Outbreaks of the twospotted spider mite on hop plants in Oregon and Washington are diminished by timing sulfur fungicides applications to periods of low spider mite abundance and slower host development (early to mid-spring). Woods 2012
The genomic divergence between Ostrinia nubilalis, a maize pest, and O. scapulalis, a hop and mugwort pest, in populations in France was analyzed. Midamegbe 2011
The genetic and pathogenic relatedness of Pseudoperonospora humuli and P. cubensis, the causal agents of downy mildew on hops and cucurbits, respectively, were investigated. Mitchell 2011
Treatment of hop plants with prohexadione-calcium during cone maturation induced increases in cone biomass and the terpenophenolic content. Kavalier 2011
The ecology and recent evolutionary history of Ostrinia nubilalis, a common hops pathogen, is studied. Pélissié 2010
Molecular variation in hop mosaic virus isolates was analyzed. Poke 2010
Application of different fungicides on hop plants in Oregon and Washington lead to dramatic changes in the size of the populations of twospotted spider mite and hop aphid, the two major arthropod pests of hops in the Northern Hemisphere, depending on the fungicide. Gent 2009
Costs associated with downy mildew control vs. crop loss in the maritime hop plantations in western Oregon were minimized by the application of the infection risk prediction models based on weather factors over 24- and 48-h periods. Gent 2009
Hop stunt viroid undergoes adaptation in cultivated host plants, which may result in the appearance of hop-pathogenic variants. Kawaguchi-Ito 2009
Gene expression in Fusarium graminearum, a fungal hop pathogen, grown on media containing hop cell wall was analyzed. Carapito 2008
The genetic diversity of apple fruit crinkle viroid, a hop pathogen, in the Akita and Yamagata Japan prefectures was analyzed. Sano 2008
Organic farming conditions had a marginal effect on the levels of alpha-acids, beta-acids, desmethylxanthohumol, and xanthohumol in only 1 of the 3 hop varieties tested. The effects of climatic and environmental conditions were more significant. Keukeleire 2007
The study focuses on the ecology and development of the butterfly Polygonia c-album, which used to have hops as the preferred plant host, in relation to the changes it has undergone due to the recent observed climate changes. Braschler 2007
Hop plants were sprayed with (Z)-jasmone attracted more damson-hop aphids during their spring migration. Pope 2007
Weed species on hop fields were identified as potential hosts for hop stunt viroid. Matousek 2007
Gold binding by native and chemically modified hops biomasses was studied for potential application for extraction of metals from industrial effluents. López 2005
The exoproteome of the fungus Fusarium graminearum grown on glucose and on hop (Humulus lupulus, L.) cell wall has been investigated. Phalip 2005
Filamentous fungi were collected from diseased hop (Humulus lupulus; L.) and DNA was prepared from 19 isolates & 7 different genera were found among the 19 isolates. Phalip 2004
The molecular variances within groups of cultivated and wild hops were homogeneous, suggesting that a similar level of molecular variability is found in both groups of accessions. Jakse 2004
It is shown that maize, mugwort, and hop host races of O. nubilalis differ not only in their host plant but also in the sex pheromone they use, because mugwort and hop are putative ancestral host plants. Pelozuelo 2004
Volatiles emitted by Tetranychus urticae-infested plants of 11 species including hop (Humulus lupulus), grapevine and ginkgo were compared. van den Boom 2004
Sixteen species of native detector plants including Humulus lupulus and Parthenocissus quinquefolia for ambient ozone have been identified for use in Central and Eastern Europe. Manning 2004
Early development of the secretory cavity of chemically fixed peltate glands in Humulus lupulus L. showed secretions with different densities, light, gray and dark, in the cytoplasm of disc cells and in the periplasmic space adjacent to the developing secretory cavity. Kim 2000
The sequence of histological and histochemical events occurring during organogenesis from Humulus lupulus var. Nugget internode-derived nodules was studied. Fortes 2000
Apple mosaic virus was found in 13 cultivars of hops Petrzik 1997
Single strand viroids infect hops, burdock and other plants causing leaf discoloration and curling Smarda 1987
The effects of the gibberellic acid treatment on cone weight, cone number, time to ripening, alpha-acid content, and organoleptic properties in three different hop cultivars ('Saazer,' 'Hüller Anfang,' and 'Hallertauer') were examined. [Article in German] Zattler 1968
A statistical model for powdery mildew of hop disease spread was developed, based on a census of commercial hop yards conducted in western Oregon in 2014-2017. Gent 2019
History of Record
November 1999
MAJOR REVISION BY: Michael C. Tims, PhD. candidate
June 2001
March 2023